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The fuel heater plug is pretty hard to miss. On the bottom rear of the fuel bowl just to the driver's side.
Besides the fuel heater, that fuse powers the PCM and IDM power relay coils (thru the PCM diode), and the wastegate control solenoid.
I'm guessing that your first instance of it popping was because a wire had shorted against ground and when you unplugged the fuel heater connector, you "fixed" that. The fact that you were able to plug the fuel heater back in makes me suspect this, and that when you plugged it back in, the harness was relocated just a bit and didn't contact that ground point till now.
Either that or you have a shorted PCM or IDM power relay, which I doubt. I can't see how one of those could short the coil to ground.
Good luck.
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First truck -- 1929 Model A Ford pickup, restored from ground up. Wish I still had it!
99.5 F250 PSD Supercab LB 4x4, ZF-6 w/SB Con OFE, 3.73LS, Boost & pyro gauges, Swamps S175/146 injectors, DP 80 HP Econo PCM (classic version  ), AIS, coolant filter w/"hokum" bracket, regulated return, heated mirror mod, lighted cupholder, Marinco heater plug-in.
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